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vascular plants







The extensive coastal, montane and woodland habitats on Thásos presumably support a rich and interesting lichen flora, but it is difficult to find any information on this group of plants for Thásos. For instance, there are no records for Thásos in Rechinger, K.H. (1943) Flora Aegea. Flora der Inseln und Halbinseln des Ägäischen Meeres. Akad. Wiss. Wein, Math.-Naturwiss. Kl., Denkschr., 105 (1). The only scientific paper to deal substantially with lichens on  Thásos is a brief study by Popnikolov (1956), which lists 27 taxa recorded from the vicinity of Thassos Town in the 1940s and I am most grateful to Andrey Kovatchev for providing a copy of this reference.  Other than this, Geisller (1977) reports six species from the island, including several from Ipsarion.

Lobaria pulmonariaThe herbarium of the Dept. of Botany at the University of Essen, Germany, contains an example of the beard lichen Evernia prunastri (L.) Ach. collected from the slopes of Profitis Ilias in 1994. I have collected a specimen of the lungwort Lobaria pulmonaria (L.) Hoffm. [det. R.G. Woods] from ancient plane trees in the hills above Thassos Town, alongside a tributary of the main river which runs past Ag. Marina. Lungworts are species of clean, moist air and may be indicative of the presence of other important epiphytic lichen species in this beautiful fragment of deciduous woodland.

Until recently this amounted to the sum of our knowledge on the lichen flora of Thásos, but further records have come to light in the course of B.F.M. Abbott's dedicated research whilst compiling a checklist of the lichens of Greece (Abbot 2009). A significant addition comes from the nineteen species recorded by I. Pišút during a visit to the island in August 2007 and two additions to the island checklist come from specimens in the herbarium of M.R.D. Seaward.

In total, Abbott (2009) lists 47 taxa from Thásos, though he notes that Popnikolov's record of Flavocetraria nivalis is almost certainly erroneous as this is an arctic-alpine species not recorded elsewhere in Greece. In 2010 Sipman & Ahti (2011) collected specimens of Cladonia from the edge of the pine forest above Maries and reported seven species new to Greece, including one, C. graeca, new to science.  Five additional species, collected by H. Sipman & T. Raus in the uplands around Toumba in 2010, are reported by Malíček, J. et al (2021). Three species of Caloplaca new to Thásos are depicted by Schumm, F. & Aptroot, A. (2019).  Other additional records appear in papers listed below and, along with the record of Lobaria pulmonaria mentioned above, this means that 75 taxa have been reliably recorded from the island. Considering the degree of attention paid to lichens in Greece generally, including the islands, this is a reasonable tally. However, over 1400 Greek taxa are documented by Abbott and some of the larger islands have over 200 taxa recorded so there are likely to be many more species awaiting discovery on Thásos.

I'm indebted to Bernard Abbott for his kindness in allowing me to consult his remarkable checklist. The records are displayed as an annually updated .pdf file, which is downloadable from the Lichens of Greece website. This site also includes a detailed and comprehensive key to all of the lichen species known from Greece, incorporating taxa from neighbouring areas that may be expected to occur in Greece. This is also freely available in .pdf format from Photographs of many Greek lichen species can be found on H. Sipman's Aegean Lichens website, though this may not be maintained indefinitely.




Acarospora versicolor Bagl. & Car. (1863)
Bagliettoa calciseda (DC.) Gueidan & Cl. Roux (2007)
Bryoria fuscescens (Gyeln.) Brodo & D. Hawksw. (1977)
Caloplaca calcitrapa Nav.-Ros., Gaya & Cl. Roux (2000)
Caloplaca crenularia (With.) J. R. Laundon (1984)
Caloplaca flavescens (Huds.) J. R. Laundon (1984)
Caloplaca hungarica H. Magn. (1944)
Caloplaca inconnexa (Nyl.) Zahlbr. (1931)
Caloplaca pollinii (A. Massal.) Jatta (1900)
Caloplaca saxicola (Hoffm.) Nordin (1972)
Caloplaca thracopontica Vondrák & Šoun (2008)
Circinaria calcarea (L.) A. Nordin, S. Savić & Tibell (2010)
Cladonia chlorophaea (Flörke ex Sommerf.) Spreng. (1827)
Cladonia ciliata var. tenuis (Flörke) Nimis (1993)
Cladonia coccifera (L.) Willd. (1787)
Cladonia coniocraea (Flörke) Spreng. (1827)
Cladonia crispata var. cetrariiformis (Delise) Vain. (1886)
Cladonia fimbriata (L.) Fr. (1831)
Cladonia floerkeana (Fr.) Flörke (1828)
Cladonia foliacea (Huds.) Willd. (1787)
Cladonia furcata (Huds.) Baumg. (1790)
Cladonia glauca Flörke (1828)
Cladonia graeca Sipman & Ahti (2011)
Cladonia humilis (With.) J. R. Laundon (1984)
Cladonia monomorpha Aptroot, Sipman & van Herk (2001)
Cladonia parasitica (Hoffm.) Hoffm. (1796)
Cladonia pyxidata (L.) Hoffm. (1796)
Cladonia ramulosa (With.) J. R. Laundon (1984)
Cladonia rangiformis Hoffm. (1796)
Cladonia squamosa Hoffm. (1796)
Cladonia subturgida Samp. (1918)
Cladonia uncialis subsp. biuncialis (Hoffm.) M. Choisy (1951)
Collema nigrescens (Huds.) DC. (1805)
Diploschistes muscorum (Scop.) R. Sant. (1980)
Diploschistes ocellatus (Vill.) Norman (1852)
Enchylium polycarpon subsp. corcyrense (Arnold) ined.
Evernia prunastri (L.) Ach. (1810)
        [Flavocetraria nivalis (L.) Kärnefelt & A. Thell (1994) - presumed incorrect]
Hypogymnia physodes (L.) Nyl. (1896)
Hypogymnia tubulosa (Schaer.) Hav. (1918)
Lathagrium cristatum (L.) Otálora, P. M. Jřrg. & Wedin (2013)
Lecanora rupicola subsp. sulphurata (Nyl.) Leuckert & Poelt (1989)
Lepra albescens (Huds.) Hafellner (2016)
Lepraria incana (L.) Ach. (1803)
Lepraria nylanderiana Kümmerl. & Leuckert (1995)
Leprocaulon microscopicum (Vill.) Gams ex D. Hawksw. (1974)
Lobaria pulmonaria (L.) Hoffm. (1796)
Lobothallia farinosa (Flörke) A. Nordin, S. Savić & Tibell (2010)
Lobothallia radiosa (Hoffm.) Hafellner (1991)
Myriolecis dispersa (Pers.) Sliwa, Z. Xin & Lumbsch (2016)
Neofuscelia pulla (Sibth.) Essl. (1978)
Ochrolechia androgyna (Hoffm.) Arnold (1885)
Ochrolechia parella (L.) A. Massal. (1852)
Parmelina tiliacea (Hoffm.) Hale (1974)
Pertusaria amarescens Nyl. (1874)
Pertusaria flavida (DC.) J. R. Laundon (1963)
Pertusaria hymenea (Ach.) Schaer. (1836)
Pertusaria pertusa (L.) Tuck. (1845)
Physconia distorta (With.) J. R. Laundon (1984)
Platismatia glauca (L.) W. L. Culb. & C. F. Culb. (1968)
Pleurosticta koflerae (Clauzade & Poelt) Elix & Lumbsch (1988)
Protoblastenia rupestris (Scop.) J. Steiner (1911)
Protoparmeliopsis vaenskaei (Cl. Roux & C. Coste) Cl. Roux (2016)
Pseudevernia furfuracea (L.) Zopf (1903)
Pseudevernia furfuracea var. ceratea (Ach.) D. Hawksw. (1969)
Ramalina carpatica Körb. (1870)
Ramalina elegans (Bagl. & Car.) Stizenb. (1891)
Ramalina farinacea (L.) Ach. (1810)
Ramalina subfarinacea (Nyl. ex Cromb.) Nyl. (1873)
Schismatomma pericleum (Ach.) Th. Fr. (1861)
Scytinium lichenoides (L.) Otálora, P. M. Jřrg. & Wedin (2013)
Solenopsora candicans (Dicks.) J. Steiner (1915)
Squamarina gypsacea (Sm.) Poelt in Kirk, P. (2020)
Thalloidima sedifolium (Scop.) Kistenich, Timdal, Bendiksby & S. Ekman (2018)
Verrucaria nigrescens Pers. (1795)
Xanthoparmelia conspersa (Ehrh. ex Ach.) Hale (1974)

Saxicolous lichens - Ipsarion summit (Oct 2001)


Abbott, B.F.M. 2009.  Checklist of the lichens and lichenicolous fungi of Greece.  Bibliotheca Lichenologica 103:1-368

Burgaz, A. R., Gutiérrez-Larruga, B., Rodríguez-Arribas, C. & Pino-Bodas, R. 2020. The genus Cladonia from Greece. Herzogia, 33: 341-376

Geissler, P. 1977. Zur Moos- und Flechtenflora Nordgriechenlands.  Bauhinia  6 (1):189-213

Knudsen, K. et al. 2015. Acarospora cinerascens (Acarosporaceae), a poorly known species from the southern Central Alps (Italy and Switzerland). Herzogia, 29: 690-697

Linda in Arcadia. 2020. Atlas of Greek lichens (and lichenicolous fungi). Privately produced.

Lindqvist, M. 1983. Grekiska lavar - ekologi och utbredning. 34 pp., Avd. för systematisk botanik, Göteborgs universitet.

Malíček, J. et al. 2021. New country records of lichenized and non-lichenized fungi from southeastern Europe. Herzogia, 34: 38-54

Pino-Bodas, al. 2015. The phenotypic features used for distinguishing species within the Cladonia furcata complex are highly homoplasious. Lichenologist 47: 287-303

Popnikolov, A. 1956. Flechten von Ost - (Griechisch) Mazedonien. Bull. de L'Institut Botanique, Academie Bulgare des Sciences, 4: 353-356. [in Bulgarian]


Schumm, F. & Aptroot, A. 2019.  Images of the lichen genus Caloplaca.  4 Volumes.  Norderstedt.


Sipman, H.J.M. & Ahti, T. 2011. Contribution to the Cladonia mycota of Greece, including Cladonia graeca sp. nov. Mycosystema, 30: 877-880


Zhang, Y. Y. et al. 2020. Squamarina (lichenised fungi) species described from China belong to at least three unrelated genera. Mycokeys, 66: 135-157